Distinguished from all other congeners by the combination of the following features: anterior portion of trunk slightly deeper than wide, jaws short, snout blunt, tip of anal fin rounded in male, caudal fin rounded in male, pelvic-fin tip reaching the anterior portion of the anal-fin base in male, dorsal-fin origin on vertical between base of 9th and 10th anal-fin rays, dorsal-fin rays 7-8, anal-fin rays 11-13, frontal squamation E-patterned, frontal scales arranged circularly, canal preopercular short and opened, contact organs on flank scales in male, longitudinal series of scales 29-32, gill rakers of first branchial arch 1 + 7, longitudinal rows of reddish brown dots stripes on flank in male, jaws black, and round black spot on dorsal portion of caudal-fin in female.
Distinguished from all other congeners by the combination of the following features: anterior portion of trunk slightly deeper than wide, jaws short, snout blunt, tip of anal fin pointed in male, caudal fin rounded in male, pelvic-fin tip reaching urogenital papilla in male, dorsal-fin origin on vertical between base of 6th and 8th anal-fin rays, dorsal-fin rays 7-8, anal-fin rays 9-10, frontal squamation S-patterned, frontal scales arranged transversely, canal preopercular absent, contact organs absent, longitudinal series of scales 29-30, gill rakers of first branchial arch 1 + 6, longitudinal rows of red dots on flank in male, transverse dark brown bar through the chin, and no distinctive black spot on dorsal portion of caudal-fin.
Distinguished from all other congeners by the combination of the following features: anterior portion of trunk slightly wider than deep, jaws short, snout blunt, tip of anal fin rounded in male, caudal fin rounded in male, pelvic-fin tip reaching anterior portion of anal-fin base in male, dorsal-fin origin on vertical between base of 8th and 9th anal-fin rays, dorsal-fin rays 6-7, anal-fin rays 9-10, frontal squamation S-patterned, frontal scales arranged transversely, canal preopercular absent, contact organs on flank scales in male, longitudinal series of scales 32, gill rakers of first branchial arch 1 + 7, transverse black bar through chin, and no black spot on dorsal portion of caudal-fin.
Distinguished from all other congeners by the combination of the following features: anterior portion of trunk slightly deeper than wide, jaws short, snout blunt, tip of anal fin slightly pointed in male, caudal fin rounded in male, pelvic-fin tip reaching middle of anal-fin base in male, dorsal-fin origin on vertical between base of 8th and 9th anal-fin rays, dorsal-fin rays 7-8, anal-fin rays 11-12, frontal squamation S-patterned, frontal scales arranged transversely, canal preopercular absent, contact organs on flank scales in male, longitudinal series of scales 32-33, gill rakers of first branchial arch 1 + 7-8, oblique rows of red dots on flank in male, transverse dark brown bar through the chin, and dark grey to black spot on dorsal portion of caudal-fin in both sexes.
Distinguished from all other congeners by the combination of the following features: anterior portion of trunk slightly deeper than wide, jaws short, snout blunt, tip of anal fin rounded in male, caudal fin truncate in male, pelvic-fin tip reaching middle of anal-fin base in male, dorsal-fin origin on vertical between base of 9th and 10th anal-fin rays, dorsal-fin rays 8-10, anal-fin rays 12-15, frontal squamation S-patterned, frontal scales arranged transversely, canal preopercular absent, contact organs on flank scales in male, longitudinal series of scales 37-41, gill rakers of first branchial arch 1 + 7, red stripes on flank, transverse black bar through the chin, and vertically elongated dark grey to black spot on dorsal portion of caudal-fin in both sexes.
Distinguished from all other congeners by the combination of the following features: anterior portion of trunk deeper than wide, jaws short, snout blunt, tip of anal fin slightly pointed in male, caudal fin rounded in male, pelvic-fin tip reaching anterior portion of anal-fin base in male, dorsal-fin origin on vertical between base of 8th and 9th anal-fin rays, dorsal-fin rays 10-11, anal-fin rays 13-14, frontal squamation F-patterned, frontal scales arranged circularly, canal preopercular absent, contact organs absent, longitudinal series of scales 31-33, gill rakers of first branchial arch 1 + 7-8, red marks of flank absent, lower jaw black, and round black spot on dorsal portion of caudal-fin in female.
